The forward clutch, which is responsible for giving you first gear in Drive, is applied and the low roller clutch assembly is holding...Upon the shift to 2nd gear, the intermediate clutch comes on, the intermediate sprag begins holding and the low roller clutch releases and begins to free-wheel. The forward clutch stays on at all times until you stop and shift into P, R or N.
So if it's binding and releasing intermittently, it sounds like something could be going wrong with the low roller clutch assembly (it's binding, debris/metal is getting into it, etc). It's also possible that there's a cross leak in the L/R circuit, partially applying the band as if it was in manual low when you're in second gear for some reason but that's a guess...
I'd recommend you replace all three one way clutch assemblies in the transmission; replace the low roller clutch assembly out of an abundance caution even if you replaced it initially as they are cheap and it's good insurance. Also pay close attn to the valve body to case gasket and valve body-spacer plate gasket for any restrictions, gasket blocking holes or tears in the gasket...
What model year is your 4L80E?